SUBGENUS Pleurothallis SECTION Macrophyllae-Fasciculatae Lindl. 1859

Characterized by the single flowered inflorescence with the peduncle hidden within a spathe, the usually short pedicels, the entire, fringed to pubescent petals that are not sigmoid in outline and a thick, obtuse, oblong to transverse lip and a shorter, broader column with an apical anther and stigma that is almost always bilobed. This subsection and amphygiae are not distinctly separated in various borderline species and for this reason they later were separated by Luer into the genus Acronia [2005] with three sections, Acronia, Amphygia and Macrophyllae-fasciculatae. TYPE Species for the SECTION

!Pleurothallis calceolaris Rchb.f. 1876

1. Pleurothallis acestrophylla Luer 1975 Ecuador and Peru

2. Pleurothallis acutilabia Luer 1999 Colombia

3. Pleurothallis adeleae Luer 1981 Ecuador and Ecuador

4. Pleurothallis adelphe Luer & Hirtz 1996 Ecuador

5. Pleurothallis adonis Luer 1976 Colombia and Ecuador

6. Pleurothallis adventurae Karremans & Bogarín 2011 Costa Rica

7. Pleurothallis aguirrei Luer & R.Escobar 2003 Colombia

8. Pleurothallis alopex Luer 1976 Bolivia and Paraguay

9. Pleurothallis altimonile Luer & R.Escobar 1981 Colombia

10. Pleurothallis ambyx Luer & R.Escobar 1981 Colombia and Ecuador

11. Pleurothallis andreaskayi Mark Wilson & B.T. Larsen 2022 Colombia, Ecuador

12. Pleurothallis andreettae (Luer) J.M.H.Shaw 2014 Ecuador

13. Pleurothallis angusta Ames & C.Schweinf. 1925 Costa Rica and Panama

14. Pleurothallis angustissima (Luer & Hirtz) J.M.H.Shaw 2014 Ecuador

15. Pleurothallis anthurioides A.Doucette 2016 Panama

16. Pleurothallis apopsis Luer 1979 Ecuador

17. Pleurothallis applanata Luer & Dalström 1996 Ecuador

18. Pleurothallis arctata Luer 1997 Peru

19. Pleurothallis ascera Luer & R. Escobar 1996 Costa Rica, Panama and Colombia

20. Pleurothallis asplundii Luer 1996 Ecuador

21. Pleurothallis aurita Schweinf 1940 Costa Rica

22. Pleurothallis baezensis (Luer & Hirtz) J.M.H.Shaw 2014 Ecuador

23. Pleurothallis barrowii Schuit. 2018 Ecuador?

24. Pleurothallis batrachus Luer & Hirtz 1996 Ecuador

25. Pleurothallis baudoensis Luer & R.Escobar 1998 Colombia

26. Pleurothallis belocardia Schltr. 1924 Colombia

27. Pleurothallis bilobulata M.M.Jiménez, Ocupa & Vélez-Abarca 2021 Ecuador

28. Pleurothallis bitumida Luer 1996 Costa Rica

29. Pleurothallis bivalvis Lindl. 1846 Ecuador, Colombia and Venezuela

30. Pleurothallis blepharopetala Schltr.1915 Ecuador

31. Pleurothallis bogarinii Pupulin & J.D. Zúñiga 2007 Costa Rica

32. Pleurothallis bothros Luer 1996 Costa Rica

33. Pleurothallis bovilingua Luer & R.Escobar 1998 Colombia

34. Pleurothallis braidiana [Luer] in ed. without location

35. Pleurothallis bucculenta Luer 1976 Ecuador

36. Pleurothallis bulbosa [Luer & Escobar] Pfahl 2012 Colombia

37. Pleurothallis calamifolia Luer & R.Escobar 1996 Colombia

38. !Pleurothallis calceolaris Rchb.f. 1876 Colombia

39. Pleurothallis callifera C.Schweinf. 1951 Venezuela

40. Pleurothallis calogramma Luer 2003 Colombia

41. Pleurothallis callosa Diaz & Pupulin 2021 Costa Rica

42. Pleurothallis calolalax Luer & R.Escobar 1998 Colombia

43. Pleurothallis canaligera Rchb.f 1877 Colombia, Ecuador and Peru

44. Pleurothallis canidentis Luer & R. Escobar 1998 Costa Rica, Panama and Colombia

45. Pleurothallis cardiochila L.O. Williams 1942 Costa Rica and Panama

46. Pleurothallis cardiocrepis Rchb.f.1888 Peru?

47. Pleurothallis cardiophylax Rchb.f. 1888 Colombia, Ecuador and Peru

48. Pleurothallis cardiostola Rchb. f. 1854 Colombia, Venezuela, Peru, Bolivia and Ecuador

49. Pleurothallis cardiothallis Rchb.f 1857 Mexico, Belize, Costa Rica, Nicaragua, Colombia and Ecuador

50. Pleurothallis carduela (Luer) J.M.H.Shaw 2014 Ecuador

51. Pleurothallis carmensotoana Mark Wilson & B.T. Larsen 2022 Ecuador, Peru

52. Pleurothallis carnosa Braid 1924 Colombia

53. Pleurothallis carrenoi Carnevali & I.Ramírez 1987 Venezuela

54. Pleurothallis cassidata Luer & R.Escobar 1998 Colombia

55. Pleurothallis castanea Mark Wilson, G.Merino & J.D.Werner 2016 Ecuador

56. Pleurothallis caucensis Mark Wilson 2017 Colombia

57. Pleurothallis cauda-hirundinis Luer & J.Portilla 2002 Ecuador

58. Pleurothallis cedrinorum Luer & Dalström 1996 Ecuador

59. Pleurothallis celsia Gal.-Tar. & J.S.Moreno 2023 Colombia

60. Pleurothallis chama Luer 1977 Ecuador

61. Pleurothallis chaparensis (Luer & R.Vásquez) J.M.H.Shaw 2014 Bolivia

62. Pleurothallis chavezii Luer 1996 Costa Rica

63. Pleurothallis chicalensis M.M.Jiménez & Baquero 2018 Ecuador

64. Pleurothallis cobriformis L.O.Williams 1940 Panama

65. Pleurothallis complanata Luer & Hirtz 1996 Ecuador

66. Pleurothallis compressa Luer 1996 Costa Rica

67. Pleurothallis conformalis Luer & Dalström 2003 Ecuador

68. Pleurothallis conicostigma Luer 1975 Ecuador

69. Pleurothallis constricta Luer & R.Escobar R. 1998 Colombia

70. Pleurothallis convexa Luer & Hirtz 1996 Ecuador

71. Pleurothallis cordata (Ruiz & Pav.) Lindl. 1830 Colombia, Venezuela, Ecuador, Peru and Bolivia

72. Pleurothallis cordifolia Rchb.f. & Wagener 1854 Costa Rica, Panama, Venzuela, Colombia and Ecuador

73. Pleurothallis coriacardia Rchb. f. 1854 Ecuador and Peru

74. Pleurothallis correllii Luer 1976 Guatemala, El Salvador and Mexico

75. Pleurothallis corysta Luer 1978 Ecuador

76. Pleurothallis crateriformis C.Schweinf. 1951 Ecuador and Peru

77. Pleurothallis crossota Luer & Dalström 1996 Ecuador

78. Pleurothallis crucifera Luer & Hirtz 1996 Ecuador

79. Pleurothallis culpameae (Luer) J.M.H.Shaw 2014 Ecuador

80. Pleurothallis cuzcoensis [Luer] ined. Peru

81. Pleurothallis cutucuensis Luer 1977 Ecuador

82. Pleurothallis cyanea Luer & R. Escobar 1984 Colombia

83. Pleurothallis dariensis (Kolan. & Szlach.) Bogarín 2014 Panama

84. Pleurothallis deflexa Luer 1975 Ecuador

85. Pleurothallis dejavu Luer & Hirtz 1988Ecuador

86. Pleurothallis depressa Luer & R. Escobar 1996 Colombia

87. Pleurothallis dewildei Luer & R.Escobar 1998 Colombia

88. Pleurothallis diabolica Luer & R. Escobar 1981 Colombia Ecuador

89. Pleurothallis diazii (Luer & Endara) J.M.H.Shaw 2014 Peru

90. Pleurothallis dibolia Luer 1975 Ecuador

91. Pleurothallis dilemma Luer 2001 Ecuador

92. Pleurothallis discoidea Lindl. 1835 Trinidad, Venezuela, Surinam, French Guiana, Ecuador and Peru

93. Pleurothallis dorotheae Luer 1980 Costa Rica

94. Pleurothallis dracuncula (Luer & Hirtz) J.M.H.Shaw 2014 Ecuador

95. Pleurothallis drewii Luer 2000

    Ecuador

96. Pleurothallis dubbeldamii Luer 2003 No Collection Data

97. Pleurothallis ensata Luer 1975 Ecuador

98. Pleurothallis epiglottis Luerb 1980 Ecuador

99. Pleurothallis erythrium Luer 1978 Ecuador

100. Pleurothallis escobarii (Luer) J.M.H.Shaw 2014 Colombia

101. Pleurothallis excavata Schltr. 1923 Costa Rica and Panama

102. Pleurothallis fantastica Ames 1936 Costa Rica

103. Pleurothallis folsomii (Luer & Endara) J.M.H.Shaw 2014 Panama

104. Pleurothallis fonnegrae (Luer & Endara) J.M.H.Shaw 2014 Colombia

105. Pleurothallis fossulata Luer & R.Escobar 1998 Colombia and Ecuador

106. Pleurothallis franciana Sierra-Aziz 2023 Colombia

107. Pleurothallis galerita Luer 1981 Ecuador

108. Pleurothallis ganymedes Luer & R.Escobar 1981 Colombia

109. Pleurothallis gargantua Luer 1996 Ecuador

110. Pleurothallis geographica Luer 1977 Ecuador

111. Pleurothallis gigiportillae A.Doucette & J.Portilla 2016 Ecuador

112. Pleurothallis giraffa Luer 1979 Ecuador

113. Pleurothallis giraldoi Luer 1999 Colombia

114. Pleurothallis glabra Luer & R.Escobar 1996 Colombia

115. Pleurothallis globosa Luer & R.Escobar 1996 Colombia

116. Pleurothallis gonzaleziorum Pupulin, Diaz & Pridgeon 2021 Costa Rica

117. Pleurothallis gonzaloi J.S. Moreno, Rinc.-González & Gal.-Tar 2022 Colombia

118. Pleurothallis grandiflora Lindl. 1835 Venezuela, Colombia, Ecuador, Peru and Bolivia

119. Pleurothallis grandilingua Pupuplin, M Diaz and Pridgeon 2021 Costa Rica

120. Pleurothallis grobleri (Luer & Hirtz) J.M.H.Shaw 2014 Ecuador

121. Pleurothallis hemileuca Luer 1982 Panama

122. Pleurothallis hoeijeri Luer & Hirtz 1988 Ecuador

123. Pleurothallis homalantha Schltr.1923 El Salvador, Honduras, Nicaragua, Costa Rica, Panama and Brazil

124. Pleurothallis imperialis Luer 1975 Ecuador and Colombia

125. Pleurothallis index Luer 1980 Ecuador and Colombia

126. Pleurothallis isthmica Luer 1977 Honduras, Costa Rica and Panama

127. Pleurothallis jaramilloi Luer 2001 Colombia

128. Pleurothallis jupiter Luer 1975 Ecuador

129. Pleurothallis kashi-menkakarai Mashendo-Jimbicti, Vélez-Abarca & M.M.Jiménez 2023 Ecuador

130. Pleurothallis killipii Garay 1956 Colombia

131. Pleurothallis lacera Luer 1975 Colombia and Ecuador

132. Pleurothallis lanigera Luer & Hirtz 1988 Ecuador

133. Pleurothallis lapoi M.M.Jiménez & Vélez-Abarca 2023 Ecuador

134. Pleurothallis latipetala Garay 1952 Colombia

135. Pleurothallis lemniscifolia Luer 1975 Ecuador

136. Pleurothallis leucantha Schltr. 1912 Guatemala

137. Pleurothallis lilijae Foldats 1968 Venezuela, Colombia and Ecuador

138. Pleurothallis linguifera Lindl. 1859 Colombia, Ecuador, Peru, Bolivia and Venezuela

139. Pleurothallis llanganatensis (Luer & Hirtz) J.M.H.Shaw 2014

140. Pleurothallis longipetala Bogarin & Belfort 2021 Costa Rica

141. Pleurothallis loreae Carnevali & I.Ramírez 2003 Venezuela

142. Pleurothallis luna-crescens Pupulin, J.Aguilar & Mel.Fernández 2017 Costa Rica

143. Pleurothallis lunaris Luer & R.Escobar 1981 Colombia

144. Pleurothallis lynniana [Luer] Comb. novo. Pfahl 2012 Ecuador

145. Pleurothallis macra Lindl. 1859 Ecuador and Peru

146. Pleurothallis macrocardia Rchb.f. 1855 Colombia, Ecuador and Bolivia

147. Pleurothallis maduroi Luer 1997 Panama

148. Pleurothallis magnipetala C.Schweinf. 1951 Peru

149. Pleurothallis mahechae J.S. Moreno, Sierra-Ariza & L.C. Pina 2022 Colombia

150. Pleurothallis marioandresavilae Vélez-Abarca & M.M.Jiménez 2023 Ecuador, Peru

151. Pleurothallis marioi Zambrano & Solano 2017 Ecuador

152. Pleurothallis marioportillae A.Doucette, H.Medina & J.Portilla 2025 Ecuador

153. Pleurothallis marthae Luer & R. Escobar 1996 Colombia

154. Pleurothallis mastodon Luer & Hirtz 1996 Ecuador

155. Pleurothallis matudana C. Schweinf. 1938 Mexico, Guatemala, Honduras, Nicaragua, Ecuador and Bolivia

156. Pleurothallis megaglossa Luer & Dalström 1996 Ecuador

157. Pleurothallis megalorhina Luer & R.Escobar 1996 Colombia

158. Pleurothallis mesopotamica Bogarin & Belfort 2021 Costa Rica

159. Pleurothallis microcardia Rchb.f. 1855 Colombia, Venezuela, Ecuador, Peru and Bolivia

160. Pleurothallis millei Schltr. 1915 Ecuador

161. Pleurothallis miniatura (Luer, Thoerle & F.Werner) J.M.H.Shaw 2014 Ecuador

162. Pleurothallis minutilabia Mark Wilson, Tobar & Á.J.Pérez 2018 Ecuador

163. Pleurothallis muriculata Luer & Hirtz 1996 Ecuador

164. Pleurothallis navisepala Pupulin, J.Aguilar & M.Díaz 2017 Costa Rica

165. Pleurothallis neobarbosae J.M.H.Shaw 2014 Peru

166. Pleurothallis neorinkei A.Doucette 2016 Colombia

167. Pleurothallis nephroglossa Schltr. 1921 Ecuador

168. Pleurothallis nipterophylla C.A.Luer 1976 Ecuador

169. Pleurothallis nitida Luer 1977 Panama

170. Pleurothallis nossax Luer & R. Escobar 1981 Colombia

171. Pleurothallis octavioi Luer & R.Escobar R. 1981 Colombia

172. Pleurothallis omoglossa Luer 1976 Ecuador

173. Pleurothallis oncoglossa Luer 1996 Costa Rica

174. Pleurothallis ortegae Luer & Hirtz 1996 Ecuador

175. Pleurothallis orygmoglossa (Luer & Dressler) J.M.H.Shaw 2014 Panama

176. Pleurothallis oscarii Archila & Chiron 2011 Guatemala

177. Pleurothallis oxapampae Luer 1982 Peru

178. Pleurothallis palliolata Ames 1922 Costa Rica and Panama

179. Pleurothallis pansamalae Schltr. 1912 Mexico, Guatemalla, Belize, El Salvador, Honduras and Nicaragua

180. Pleurothallis papillingua A.Doucette & J.Portilla 2016 Ecuador

181. Pleurothallis paquishae Luer 1996 Ecuador and Peru

182. Pleurothallis peculiaris Luer 1976 Panama

183. Pleurothallis pelicophora Luer 2001 Ecuador

184. Pleurothallis penelops Luer 1978 Ecuador

185. Pleurothallis perijaensis Dunst. 1978 Venezuela, Colombia and Ecuador

186. Pleurothallis peroniocephala Luer 1975 Ecuador

187. Pleurothallis perryi Luer 1979 Ecuador

188. Pleurothallis persimilis Luer & Hirtz 2003 Ecuador

189. Pleurothallis petroana Sierra-Ariza 2023 Colombia

190. Pleurothallis phyllocardia Rchb.f. 1866 Costa Rica and Panama

191. Pleurothallis phyllocardioides Schltr. 1923 Guatemala, Nicaragua, Costa Rica, Panama, Colombia, Ecuador, Peru, Bolivia and Venezuela

192. Pleurothallis phymatodea Luer 1976 Ecuador

193. Pleurothallis pileata Luer & R.Escobar 1982 Colombia

194. Pleurothallis platypetala Luer & R.Escobar 1996 Colombia

195. Pleurothallis platysepala Schltr. 1920 Colombia

196. Pleurothallis portillae Luer 1979 Colombia and Ecuador

197. Pleurothallis prolaticollaris Luer 1977 Ecuador

198. Pleurothallis pudica Pupulin, J.Aguilar & M.Díaz 2017 Costa Rica

199. Pleurothallis pyelophera [Luer] comb. novo. Pfahl 2013 Ecuador

200. Pleurothallis quitu-cara Carrera & Baquero 2018 Ecuador

201. Pleurothallis radula Luer 1979 Costa Rica

202. Pleurothallis ramiromedinae Thoerle & Hirtz 2018 Colombia, Ecuador

203. Pleurothallis rectipetala Ames & Schweinf. 1925 Panama

204. Pleurothallis recurvata Luer & Hirtz 1996 Ecuador

205. Pleurothallis reginae Garay 1953 Colombia

206. Pleurothallis reptans Luer 1981 Ecuador

207. Pleurothallis rhinocera (Luer & Sijm) J.M.H.Shaw 2013 Ecuador

208. Pleurothallis rhodoglossa Schltr.1922 Costa Rica, Panama, Colombia and Ecuador

209. Pleurothallis rikseniana Mark Wilson & B.T. Larsen 2022 ?Ecuador?

210. Pleurothallis ripleyi Luer 1979 Ecuador

211. Pleurothallis ruberrima Lindley 1846 Colombia, Ecuador, Peru, Bolivia and Venezuela

212. Pleurothallis rubrifolia Mark Wilson, Tobar & Salas Guerr. 2016 Ecuador, Peru

213. Pleurothallis rubroinversa Luer 1975 Ecuador

214. Pleurothallis rugosa Luer & R.Escobar 1982 Colombia

215. Pleurothallis sabanillae M.M.Jiménez & Vélez-Abarca 2023 Ecuador

216. Pleurothallis sagittilabia Luer 1998 Colombia

217. Pleurothallis sanchoi Ames 1923 Guatemala, Costa Rica and Panama

218. Pleurothallis sannio Luer & R.Escobar 1988 Colombia

219. Pleurothallis sarcochila Garay 1953 Ecuador

220. Pleurothallis scabrilinguis Lindl. 1836 Colombia Ecuador and Peru

221. Pleurothallis scaphipetala Luer 1996 Costa Rica

222. Pleurothallis scotinantha Pupulin, M.Díaz & J.Aguilar 2017 Costa Rica

223. Pleurothallis scurrula Luer 1976 Peru

224. Pleurothallis semiscabra Lindl. 1846 Venezuela

225. Pleurothallis sergioi Luer & R.Escobar 2003 Colombia

226. Pleurothallis serricardia Schltr. 1924 Colombia

227. Pleurothallis sigynes Luer 1976 Colombia & Ecuador

228. Pleurothallis silvae-pacis Karremans 2011 Costa Rica

229. Pleurothallis siphoglossa Luer & R. Escobar 1996 Colombia

230. Pleurothallis sotarae Schltr. 1920 Colombia

231. Pleurothallis sphaerantha Luer 1975 Ecuador

232. Pleurothallis stelidilabia Luer 1979 Ecuador

233. Pleurothallis stellata I.Ramírez & Carnevali 2003 Venezuela

234. Pleurothallis stenosepala Rolfe 1892 Venezuela, Colombia, Ecuador, Peru and Bolivia

235. Pleurothallis stenota Luer 1977 Colombia and Ecuador

236. Pleurothallis subtilis C.Schweinf. 1951 Venezuela and Colombia

237. Pleurothallis x subversa Pupulin & Bogarin 2021 Costa Rica

238. Pleurothallis tamaensis Foldats 1968 northwestern Venezuela

239. Pleurothallis tapatiensis Pupulin, M Diaz & Pridgeon 2021 Costa Rica

240. Pleurothallis teaguei Luer 1996 Ecuador

241. Pleurothallis tinajillensis M.M.Jiménez, H. Garzón & Vélez-Abarca 2023 Ecuador

242. Pleurothallis titan Luer 1977 Panama and Colombia

243. Pleurothallis tobarii [Luer & Hirtz] comb. novo. Pfahl 2012 Ecuador

244. Pleurothallis tonduzii Schltr. 1918 Costa Rica, Panama and Peru?

245. Pleurothallis trachysepala Kraenzl. 1905 Colombia, Ecuador, Peru

246. Pleurothallis tragulosa Luer & Hirtz 1996 Ecuador

247. Pleurothallis transversilabia Carnevali & I.Ramírez 2000 Venezuela

248. Pleurothallis triangulilabia C.Schweinf. 1937 Costa Rica

249. Pleurothallis tridentata Klotzsch 1840 Venezuela, Colombia, Ecuador, Peru? and Bolivia?

250. Pleurothallis troglodytes Luer 1982 Ecuador

251. Pleurothallis tyria Luer & Hirtz 1996 Ecuador

252. Pleurothallis undulata Poepp. & Endl. 1836 Peru

253. Pleurothallis upanoensis Luer & Hirtz 2003 Ecuador

254. Pleurothallis valladolidensis Luer 1983 Ecuador

255. Pleurothallis variabilis Luer 1979 Colombia and Ecuador

256. Pleurothallis villahermosae Sierra-Ariza, Rinc.-González & Villanueva 2022 Colombia

257. Pleurothallis vinealis Luer & R.Escobar 1986 Colombia and Ecuador

258. Pleurothallis volans Luer & Hirtz 1996 Ecuador

259. Pleurothallis whitteniana MWilson & B T Larsen 2022 Ecuador, Peru

260. Pleurothallis zarumae Luer & Hirtz 1996 Ecuador

     --------------------------------------------------------------------------------------------------------------------------