SUBGENUS Pleurothallis SECTION Macrophyllae-Fasciculatae Lindl. 1859

Characterized by the single flowered inflorescence with the peduncle hidden within a spathe, the usually short pedicels, the entire, fringed to pubescent petals that are not sigmoid in outline and a thick, obtuse, oblong to transverse lip and a shorter, broader column with an apical anther and stigma that is almost always bilobed. This subsection and amphygiae are not distinctly separated in various borderline species and for this reason they later were separated by Luer into the genus Acronia [2005] with three sections, Acronia, Amphygia and Macrophyllae-fasciculatae. TYPE Species for the SECTION

!Pleurothallis calceolaris Rchb.f. 1876

  1. Pleurothallis acestrophylla Luer 1975 Ecuador and Peru

  2. Pleurothallis acutilabia Luer 1999 Colombia

  3. Pleurothallis adeleae Luer 1981 Ecuador and Ecuador

  4. Pleurothallis adelphe Luer & Hirtz 1996 Ecuador

  5. Pleurothallis adonis Luer 1976 Colombia and Ecuador

  6. Pleurothallis adventurae Karremans & Bogarín 2011 Costa Rica

  7. Pleurothallis aguirrei Luer & R.Escobar 2003 Colombia

  8. Pleurothallis alopex Luer 1976 Bolivia and Paraguay

  9. Pleurothallis altimonile Luer & R.Escobar 1981 Colombia

  10. Pleurothallis ambyx Luer & R.Escobar 1981 Colombia and Ecuador

  11. Pleurothallis andreaskayi Mark Wilson & B.T. Larsen 2022 Colombia, Ecuador

  12. Pleurothallis andreettae (Luer) J.M.H.Shaw 2014 Ecuador

  13. Pleurothallis angusta Ames & C.Schweinf. 1925 Costa Rica and Panama

  14. Pleurothallis angustissima (Luer & Hirtz) J.M.H.Shaw 2014 Ecuador

  15. Pleurothallis anthurioides A.Doucette 2016 Panama

  16. Pleurothallis apopsis Luer 1979 Ecuador

  17. Pleurothallis applanata Luer & Dalström 1996 Ecuador

  18. Pleurothallis arctata Luer 1997 Peru

  19. Pleurothallis ascera Luer & R. Escobar 1996 Costa Rica, Panama and Colombia

  20. Pleurothallis asplundii Luer 1996 Ecuador

  21. Pleurothallis aurita Schweinf 1940 Costa Rica

  22. Pleurothallis austinrumleyi K.W.Holcomb 2024 Colombia

  23. Pleurothallis baezensis (Luer & Hirtz) J.M.H.Shaw 2014 Ecuador

  24. Pleurothallis barrowii Schuit. 2018 Ecuador?

  25. Pleurothallis batrachus Luer & Hirtz 1996 Ecuador

  26. Pleurothallis baudoensis Luer & R.Escobar 1998 Colombia

  27. Pleurothallis belocardia Schltr. 1924 Colombia

  28. Pleurothallis bilobulata M.M.Jiménez, Ocupa & Vélez-Abarca 2021 Ecuador

  29. Pleurothallis bitumida Luer 1996 Costa Rica

  30. Pleurothallis bivalvis Lindl. 1846 Ecuador, Colombia and Venezuela

  31. Pleurothallis blepharopetala Schltr.1915 Ecuador

  32. Pleurothallis bogarinii Pupulin & J.D. Zúñiga 2007 Costa Rica

  33. Pleurothallis bothros Luer 1996 Costa Rica

  34. Pleurothallis bovilingua Luer & R.Escobar 1998 Colombia

  35. Pleurothallis braidiana [Luer] in ed. without location

    Pleurothallis brinkmaniana K.W. Holcomb 2024 Colombia

  36. Pleurothallis bucculenta Luer 1976 Ecuador

  37. Pleurothallis bulbosa [Luer & Escobar] Pfahl 2012 Colombia

  38. Pleurothallis calamifolia Luer & R.Escobar 1996 Colombia

  39. !Pleurothallis calceolaris Rchb.f. 1876 Colombia

  40. Pleurothallis callifera C.Schweinf. 1951 Venezuela

  41. Pleurothallis calogramma Luer 2003 Colombia

  42. Pleurothallis callosa Diaz & Pupulin 2021 Costa Rica

  43. Pleurothallis calolalax Luer & R.Escobar 1998 Colombia

  44. Pleurothallis canaligera Rchb.f 1877 Colombia, Ecuador and Peru

  45. Pleurothallis canidentis Luer & R. Escobar 1998 Costa Rica, Panama and Colombia

  46. Pleurothallis cardiochila L.O. Williams 1942 Costa Rica and Panama

  47. Pleurothallis cardiocrepis Rchb.f.1888 Peru?

  48. Pleurothallis cardiophylax Rchb.f. 1888 Colombia, Ecuador and Peru

  49. Pleurothallis cardiostola Rchb. f. 1854 Colombia, Venezuela, Peru, Bolivia and Ecuador

  50. Pleurothallis cardiothallis Rchb.f 1857 Mexico, Belize, Costa Rica, Nicaragua, Colombia and Ecuador

  51. Pleurothallis carduela (Luer) J.M.H.Shaw 2014 Ecuador

  52. Pleurothallis carmensotoana Mark Wilson & B.T. Larsen 2022 Ecuador, Peru

  53. Pleurothallis carnosa Braid 1924 Colombia

  54. Pleurothallis carrenoi Carnevali & I.Ramírez 1987 Venezuela

  55. Pleurothallis cassidata Luer & R.Escobar 1998 Colombia

  56. Pleurothallis castanea Mark Wilson, G.Merino & J.D.Werner 2016 Ecuador

  57. Pleurothallis caucensis Mark Wilson 2017 Colombia

  58. Pleurothallis cauda-hirundinis Luer & J.Portilla 2002 Ecuador

  59. Pleurothallis cedrinorum Luer & Dalström 1996 Ecuador

  60. Pleurothallis celsia Gal.-Tar. & J.S.Moreno 2023 Colombia

  61. Pleurothallis chama Luer 1977 Ecuador

  62. Pleurothallis chaparensis (Luer & R.Vásquez) J.M.H.Shaw 2014 Bolivia

  63. Pleurothallis chavezii Luer 1996 Costa Rica

  64. Pleurothallis chicalensis M.M.Jiménez & Baquero 2018 Ecuador

  65. Pleurothallis cobriformis L.O.Williams 1940 Panama

  66. Pleurothallis complanata Luer & Hirtz 1996 Ecuador

  67. Pleurothallis compressa Luer 1996 Costa Rica

  68. Pleurothallis conformalis Luer & Dalström 2003 Ecuador

  69. Pleurothallis conicostigma Luer 1975 Ecuador

  70. Pleurothallis constricta Luer & R.Escobar R. 1998 Colombia

  71. Pleurothallis convexa Luer & Hirtz 1996 Ecuador

  72. Pleurothallis cordata (Ruiz & Pav.) Lindl. 1830 Colombia, Venezuela, Ecuador, Peru and Bolivia

  73. Pleurothallis cordifolia Rchb.f. & Wagener 1854 Costa Rica, Panama, Venzuela, Colombia and Ecuador

  74. Pleurothallis coriacardia Rchb. f. 1854 Ecuador and Peru

  75. Pleurothallis correllii Luer 1976 Guatemala, El Salvador and Mexico

  76. Pleurothallis corysta Luer 1978 Ecuador

  77. Pleurothallis crateriformis C.Schweinf. 1951 Ecuador and Peru

  78. Pleurothallis crossota Luer & Dalström 1996 Ecuador

  79. Pleurothallis crucifera Luer & Hirtz 1996 Ecuador

  80. Pleurothallis culpameae (Luer) J.M.H.Shaw 2014 Ecuador

  81. Pleurothallis cuzcoensis [Luer] ined. Peru

  82. Pleurothallis cutucuensis Luer 1977 Ecuador

  83. Pleurothallis cyanea Luer & R. Escobar 1984 Colombia

  84. Pleurothallis dariensis (Kolan. & Szlach.) Bogarín 2014 Panama

  85. Pleurothallis deflexa Luer 1975 Ecuador

  86. Pleurothallis dejavu Luer & Hirtz 1988Ecuador

  87. Pleurothallis depressa Luer & R. Escobar 1996 Colombia

  88. Pleurothallis dewildei Luer & R.Escobar 1998 Colombia

  89. Pleurothallis diabolica Luer & R. Escobar 1981 Colombia Ecuador

  90. Pleurothallis diazii (Luer & Endara) J.M.H.Shaw 2014 Peru

  91. Pleurothallis dibolia Luer 1975 Ecuador

  92. Pleurothallis dilemma Luer 2001 Ecuador

  93. Pleurothallis discoidea Lindl. 1835 Trinidad, Venezuela, Surinam, French Guiana, Ecuador and Peru

  94. Pleurothallis dorotheae Luer 1980 Costa Rica

  95. Pleurothallis dracuncula (Luer & Hirtz) J.M.H.Shaw 2014 Ecuador

  96. Pleurothallis drewii Luer 2000

    Ecuador

  97. Pleurothallis dubbeldamii Luer 2003 No Collection Data

  98. Pleurothallis elliottii K.W.Holcomb 2024 eCUADOR

  99. Pleurothallis ensata Luer 1975 Ecuador

  100. Pleurothallis epiglottis Luerb 1980 Ecuador

  101. Pleurothallis erythrium Luer 1978 Ecuador

  102. Pleurothallis escobarii (Luer) J.M.H.Shaw 2014 Colombia

  103. Pleurothallis excavata Schltr. 1923 Costa Rica and Panama

  104. Pleurothallis fantastica Ames 1936 Costa Rica

  105. Pleurothallis folsomii (Luer & Endara) J.M.H.Shaw 2014 Panama

  106. Pleurothallis fonnegrae (Luer & Endara) J.M.H.Shaw 2014 Colombia

  107. Pleurothallis fossulata Luer & R.Escobar 1998 Colombia and Ecuador

  108. Pleurothallis franciana Sierra-Aziz 2023 Colombia

  109. Pleurothallis galerita Luer 1981 Ecuador

  110. Pleurothallis ganymedes Luer & R.Escobar 1981 Colombia

  111. Pleurothallis gargantua Luer 1996 Ecuador

  112. Pleurothallis geographica Luer 1977 Ecuador

  113. Pleurothallis gigiportillae A.Doucette & J.Portilla 2016 Ecuador

  114. Pleurothallis giraffa Luer 1979 Ecuador

  115. Pleurothallis giraldoi Luer 1999 Colombia

  116. Pleurothallis glabra Luer & R.Escobar 1996 Colombia

  117. Pleurothallis globosa Luer & R.Escobar 1996 Colombia

  118. Pleurothallis gonzaleziorum Pupulin, Diaz & Pridgeon 2021 Costa Rica

  119. Pleurothallis gonzaloi J.S. Moreno, Rinc.-González & Gal.-Tar 2022 Colombia

  120. Pleurothallis grandiflora Lindl. 1835 Venezuela, Colombia, Ecuador, Peru and Bolivia

  121. Pleurothallis grandilingua Pupuplin, M Diaz and Pridgeon 2021 Costa Rica

  122. Pleurothallis grobleri (Luer & Hirtz) J.M.H.Shaw 2014 Ecuador

  123. Pleurothallis hemileuca Luer 1982 Panama

  124. Pleurothallis hoeijeri Luer & Hirtz 1988 Ecuador

  125. Pleurothallis homalantha Schltr.1923 El Salvador, Honduras, Nicaragua, Costa Rica, Panama and Brazil

  126. Pleurothallis imperialis Luer 1975 Ecuador and Colombia

  127. Pleurothallis index Luer 1980 Ecuador and Colombia

  128. Pleurothallis isthmica Luer 1977 Honduras, Costa Rica and Panama

  129. Pleurothallis jaramilloi Luer 2001 Colombia

  130. Pleurothallis jennydandreae K.W. Holcomb 2024 Colombia

  131. Pleurothallis josefinae K.W.Holcomb 2024 Ecuador

  132. Pleurothallis jupiter Luer 1975 Ecuador

  133. Pleurothallis kashi-menkakarai Mashendo-Jimbicti, Vélez-Abarca & M.M.Jiménez 2023 Ecuador

  134. Pleurothallis killipii Garay 1956 Colombia

  135. Pleurothallis lacera Luer 1975 Colombia and Ecuador

  136. Pleurothallis lanigera Luer & Hirtz 1988 Ecuador

  137. Pleurothallis lapoi M.M.Jiménez & Vélez-Abarca 2023 Ecuador

  138. Pleurothallis latipetala Garay 1952 Colombia

  139. Pleurothallis lemniscifolia Luer 1975 Ecuador

  140. Pleurothallis leucantha Schltr. 1912 Guatemala

  141. Pleurothallis lilijae Foldats 1968 Venezuela, Colombia and Ecuador

  142. Pleurothallis linguifera Lindl. 1859 Colombia, Ecuador, Peru, Bolivia and Venezuela

  143. Pleurothallis llanganatensis (Luer & Hirtz) J.M.H.Shaw 2014

  144. Pleurothallis longipetala Bogarin & Belfort 2021 Costa Rica

  145. Pleurothallis loreae Carnevali & I.Ramírez 2003 Venezuela

  146. Pleurothallis luna-crescens Pupulin, J.Aguilar & Mel.Fernández 2017 Costa Rica

  147. Pleurothallis lunaris Luer & R.Escobar 1981 Colombia

  148. Pleurothallis lynniana [Luer] Comb. novo. Pfahl 2012 Ecuador

  149. Pleurothallis macra Lindl. 1859 Ecuador and Peru

  150. Pleurothallis macrocardia Rchb.f. 1855 Colombia, Ecuador and Bolivia

  151. Pleurothallis maduroi Luer 1997 Panama

  152. Pleurothallis magnipetala C.Schweinf. 1951 Peru

  153. Pleurothallis mahechae J.S. Moreno, Sierra-Ariza & L.C. Pina 2022 Colombia

  154. Pleurothallis marioandresavilae Vélez-Abarca & M.M.Jiménez 2023 Ecuador, Peru

  155. Pleurothallis marioi Zambrano & Solano 2017 Ecuador

  156. Pleurothallis marioportillae A.Doucette, H.Medina & J.Portilla 2025 Ecuador

  157. Pleurothallis marthae Luer & R. Escobar 1996 Colombia

  158. Pleurothallis mastodon Luer & Hirtz 1996 Ecuador

  159. Pleurothallis matudana C. Schweinf. 1938 Mexico, Guatemala, Honduras, Nicaragua, Ecuador and Bolivia

  160. Pleurothallis megaglossa Luer & Dalström 1996 Ecuador

  161. Pleurothallis megalorhina Luer & R.Escobar 1996 Colombia

  162. Pleurothallis mesopotamica Bogarin & Belfort 2021 Costa Rica

  163. Pleurothallis microcardia Rchb.f. 1855 Colombia, Venezuela, Ecuador, Peru and Bolivia

  164. Pleurothallis millei Schltr. 1915 Ecuador

  165. Pleurothallis miniatura (Luer, Thoerle & F.Werner) J.M.H.Shaw 2014 Ecuador

  166. Pleurothallis minutilabia Mark Wilson, Tobar & Á.J.Pérez 2018 Ecuador

  167. Pleurothallis muriculata Luer & Hirtz 1996 Ecuador

  168. Pleurothallis navisepala Pupulin, J.Aguilar & M.Díaz 2017 Costa Rica

  169. Pleurothallis neobarbosae J.M.H.Shaw 2014 Peru

  170. Pleurothallis neorinkei A.Doucette 2016 Colombia

  171. Pleurothallis nephroglossa Schltr. 1921 Ecuador

  172. Pleurothallis nipterophylla C.A.Luer 1976 Ecuador

  173. Pleurothallis nitida Luer 1977 Panama

  174. Pleurothallis nossax Luer & R. Escobar 1981 Colombia

  175. Pleurothallis octavioi Luer & R.Escobar R. 1981 Colombia

  176. Pleurothallis omoglossa Luer 1976 Ecuador

  177. Pleurothallis oncoglossa Luer 1996 Costa Rica

  178. Pleurothallis ortegae Luer & Hirtz 1996 Ecuador

  179. Pleurothallis orygmoglossa (Luer & Dressler) J.M.H.Shaw 2014 Panama

  180. Pleurothallis oscarii Archila & Chiron 2011 Guatemala

  181. Pleurothallis oxapampae Luer 1982 Peru

  182. Pleurothallis palliolata Ames 1922 Costa Rica and Panama

  183. Pleurothallis pansamalae Schltr. 1912 Mexico, Guatemalla, Belize, El Salvador, Honduras and Nicaragua

  184. Pleurothallis papillingua A.Doucette & J.Portilla 2016 Ecuador

  185. Pleurothallis paquishae Luer 1996 Ecuador and Peru

  186. Pleurothallis peculiaris Luer 1976 Panama

  187. Pleurothallis pelicophora Luer 2001 Ecuador

  188. Pleurothallis penelops Luer 1978 Ecuador

  189. Pleurothallis perijaensis Dunst. 1978 Venezuela, Colombia and Ecuador

  190. Pleurothallis peroniocephala Luer 1975 Ecuador

  191. Pleurothallis perryi Luer 1979 Ecuador

  192. Pleurothallis persimilis Luer & Hirtz 2003 Ecuador

  193. Pleurothallis petroana Sierra-Ariza 2023 Colombia

  194. Pleurothallis phyllocardia Rchb.f. 1866 Costa Rica and Panama

  195. Pleurothallis phyllocardioides Schltr. 1923 Guatemala, Nicaragua, Costa Rica, Panama, Colombia, Ecuador, Peru, Bolivia and Venezuela

  196. Pleurothallis phymatodea Luer 1976 Ecuador

  197. Pleurothallis pileata Luer & R.Escobar 1982 Colombia

  198. Pleurothallis platypetala Luer & R.Escobar 1996 Colombia

  199. Pleurothallis platysepala Schltr. 1920 Colombia

  200. Pleurothallis portillae Luer 1979 Colombia and Ecuador

  201. Pleurothallis prolaticollaris Luer 1977 Ecuador

  202. Pleurothallis pudica Pupulin, J.Aguilar & M.Díaz 2017 Costa Rica

  203. Pleurothallis pyelophera [Luer] comb. novo. Pfahl 2013 Ecuador

  204. Pleurothallis quitu-cara Carrera & Baquero 2018 Ecuador

  205. Pleurothallis radula Luer 1979 Costa Rica

  206. Pleurothallis ramiromedinae Thoerle & Hirtz 2018 Colombia, Ecuador

  207. Pleurothallis rectipetala Ames & Schweinf. 1925 Panama

  208. Pleurothallis recurvata Luer & Hirtz 1996 Ecuador

  209. Pleurothallis reginae Garay 1953 Colombia

  210. Pleurothallis reptans Luer 1981 Ecuador

  211. Pleurothallis rhinocera (Luer & Sijm) J.M.H.Shaw 2013 Ecuador

  212. Pleurothallis rhodoglossa Schltr.1922 Costa Rica, Panama, Colombia and Ecuador

  213. Pleurothallis rikseniana Mark Wilson & B.T. Larsen 2022 ?Ecuador?

  214. Pleurothallis ripleyi Luer 1979 Ecuador

  215. Pleurothallis ruberrima Lindley 1846 Colombia, Ecuador, Peru, Bolivia and Venezuela

  216. Pleurothallis rubrifolia Mark Wilson, Tobar & Salas Guerr. 2016 Ecuador, Peru

  217. Pleurothallis rubroinversa Luer 1975 Ecuador

  218. Pleurothallis rugosa Luer & R.Escobar 1982 Colombia

  219. Pleurothallis sabanillae M.M.Jiménez & Vélez-Abarca 2023 Ecuador

  220. Pleurothallis sagittilabia Luer 1998 Colombia

  221. Pleurothallis sanchoi Ames 1923 Guatemala, Costa Rica and Panama

  222. Pleurothallis sannio Luer & R.Escobar 1988 Colombia

  223. Pleurothallis sarcochila Garay 1953 Ecuador

  224. Pleurothallis scabrilinguis Lindl. 1836 Colombia Ecuador and Peru

  225. Pleurothallis scaphipetala Luer 1996 Costa Rica

  226. Pleurothallis scotinantha Pupulin, M.Díaz & J.Aguilar 2017 Costa Rica

  227. Pleurothallis scurrula Luer 1976 Peru

  228. Pleurothallis semiscabra Lindl. 1846 Venezuela

  229. Pleurothallis sergioi Luer & R.Escobar 2003 Colombia

  230. Pleurothallis serricardia Schltr. 1924 Colombia

  231. Pleurothallis sigynes Luer 1976 Colombia & Ecuador

  232. Pleurothallis silvae-pacis Karremans 2011 Costa Rica

  233. Pleurothallis siphoglossa Luer & R. Escobar 1996 Colombia

  234. Pleurothallis sotarae Schltr. 1920 Colombia

  235. Pleurothallis sphaerantha Luer 1975 Ecuador

  236. Pleurothallis stelidilabia Luer 1979 Ecuador

  237. Pleurothallis stellata I.Ramírez & Carnevali 2003 Venezuela

  238. Pleurothallis stenosepala Rolfe 1892 Venezuela, Colombia, Ecuador, Peru and Bolivia

  239. Pleurothallis stenota Luer 1977 Colombia and Ecuador

  240. Pleurothallis subtilis C.Schweinf. 1951 Venezuela and Colombia

  241. Pleurothallis x subversa Pupulin & Bogarin 2021 Costa Rica

  242. Pleurothallis tamaensis Foldats 1968 northwestern Venezuela

  243. Pleurothallis tapatiensis Pupulin, M Diaz & Pridgeon 2021 Costa Rica

  244. Pleurothallis teaguei Luer 1996 Ecuador

  245. Pleurothallis tinajillensis M.M.Jiménez, H. Garzón & Vélez-Abarca 2023 Ecuador

  246. Pleurothallis titan Luer 1977 Panama and Colombia

  247. Pleurothallis tobarii [Luer & Hirtz] comb. novo. Pfahl 2012 Ecuador

  248. Pleurothallis tonduzii Schltr. 1918 Costa Rica, Panama and Peru?

  249. Pleurothallis trachysepala Kraenzl. 1905 Colombia, Ecuador, Peru

  250. Pleurothallis tragulosa Luer & Hirtz 1996 Ecuador

  251. Pleurothallis transversilabia Carnevali & I.Ramírez 2000 Venezuela

  252. Pleurothallis triangulilabia C.Schweinf. 1937 Costa Rica

  253. Pleurothallis tridentata Klotzsch 1840 Venezuela, Colombia, Ecuador, Peru? and Bolivia?

  254. Pleurothallis troglodytes Luer 1982 Ecuador

  255. Pleurothallis tyria Luer & Hirtz 1996 Ecuador

  256. Pleurothallis undulata Poepp. & Endl. 1836 Peru

  257. Pleurothallis upanoensis Luer & Hirtz 2003 Ecuador

  258. Pleurothallis valladolidensis Luer 1983 Ecuador

  259. Pleurothallis variabilis Luer 1979 Colombia and Ecuador

  260. Pleurothallis villahermosae Sierra-Ariza, Rinc.-González & Villanueva 2022 Colombia

  261. Pleurothallis vinealis Luer & R.Escobar 1986 Colombia and Ecuador

  262. Pleurothallis volans Luer & Hirtz 1996 Ecuador

  263. Pleurothallis whitteniana MWilson & B T Larsen 2022 Ecuador, Peru

  264. Pleurothallis zarumae Luer & Hirtz 1996 Ecuador

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